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You has summoned the fictional bioscience expert/genetics nerd!

Maybe the dragons can breed with humans because they created human forms, so they can copy all that? Roy has a teeny bit of ice dragon in him, or he's half ice dragon if Ninian is full dragon/dragon status is dominant, which would in turn make Roy a dragon himself!

Or, it depends on the form the mother was in when making the kid? Nergal seems like he's be into some kinky scaley shit with Aenir, resulting in Ninian and Nils hatching from eggs as little ice dragons that can take human form later. Roy was probably a live human style birth, unless Eliwood and Ninian are also kinky.

Tellius covers this better, with the branded. I'm saying laguz kids can shift from birth, they just have to figure out how. So you have beast tribe parents, it's very possible for them to go check on their kid and find a kitten the size of a human baby. Sign of a smart kid. Beast laguz are born blind and deaf too.

Fodlan crests, there can be carriers of the crest. Miklan was a carrier.

Etc.

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